CHARACTERS OF LIVING THINGS

Characteristics of Living Beings

1. Growth: All living organisms grow in size and number. Some non-living things also grow. For example mountains develop over a period of time. But growth of non-living things is a result of accumulation of substances from outside. On the other hand growth of living beings comes from inside. A continuous process of cell division makes this growth possible. Most of the plants show growth in size throughout their life. But animals grow to a certain limit. After this the cell division facilitates repair in the organism. The growth from inside is also called organic growth.

2. Reproduction: All living beings reproduce their progeny. Progeny are more or less similar to their parents. Reproduction can be sexual or asexual. Most of the unicellular and less complex organisms reproduce by asexual methods. Multicellular and complex organisms reproduce by sexual methods. Reproduction by sexual method involves formation of male and female gametes and fusion of those gametes results in formation of zygotes, which ultimately give birth to the new progeny.

3. Metabolism: All living organisms are made of chemicals. These chemicals are constantly undergoing some change inside the body of a living organism. The process of this constant change is called metabolism. Metabolism can be classified in two types:

(a) Anabolism: The process which results in synthesis of something is called anabolism. Photosynthesis is a good example of anabolism as it involves synthesis of carbohydrates.

(b) Catabolism: The process which results in breaking down of a compound is called catabolism. Respiration is a good example of catabolism as it involves breaking down of carbohydrates into carbon-dioxide and water.

4. Response to Outside Stimuli: All living organisms show a response to outside stimulus. These stimuli can be light, heat, other organisms or chemicals. For example all green plants grow in the direction from which sunlight comes. All animals are sensitive to heat.

NOMENCLATURE

Nomenclature of Living Beings Biodiversity: The planet earth is full of variety of animals and plants. Every distinct geographical location has its own set of flora and fauna. This variety of living beings found in a given geographical area is called biodiversity of that area. Nomenclature and Identification: Because of the huge diversity present in the living world it became necessary to develop a system to classify and name all plants and animals. International Code for Botanical Nomenclature has developed a system for identification and classification of plants. International Code of Zoological Nomenclature has developed a system for identifying and classifying the animals.

General Rules for Nomenclature: 1. Biological names are generally in Latin and written in italics. They are Latinised or derived from Latin irrespective of their origin. 2. The first word in a biological name represents the genus while the second component denotes the specific epithet. 3. Both the words in a biological name, when handwritten, are separately underlined, or printed in italics to indicate their Latin origin. 4. The first word denoting the genus starts with a capital letter while the specific epithet starts with a small letter.

TAXONOMY

Taxonomic Categories Classification involves hierarchy of steps in which each step represents a rank or category. Since the category is a part of overall taxonomic arrangement, it is called the taxonomic category. All categories together constitute the taxonomic hierarchy. Each category represents a rank and is commonly termed as taxon (pl.: taxa). Species A group of individual organisms with fundamental similarities is called species. Certain distinct morphological characters distinguish one species from the other closely related species. For example; Solanum tuberosum (potato),Solanum nigrum (Black nightshade) and Solanum melongena (aubergine or eggplant) are different species from the genus Solanum. Similarly; Panthera leo (lion) andPanthera tigris (tiger) are different species from the genus Panthera. Genus  A genus is a group which is comprised of closely related speciesWe can say that genera are aggregates of closely related species. For example, potato, tomato and brinjal are three different species but all belong to the genus Solanum. Similarly; Lion (Panthera leo), leopard (P. pardus) and tiger (P. tigris) belong to the genus Panthera. Family The next category, Family, has a group of related genera with still less number of similarities as compared to genus and species. Families are characterised on the basis of both vegetative and reproductive features of plant species. For example; three different genera Solanum, Petunia and Datura are placed in the family Solanaceae. Similarly; genus Panthera, comprising lion, tiger, leopard is put along with genus, Felis (cats) belong to the family Felidae.	Order  Order is the group of families which exhibit a few similar characters. The similar characters are less in number as compared to different genera included in a family. Plant families like Convolvulaceae, Solanaceae are included in the order Polymoniales on the basis of floral characters. The animal order, Carnivora, includes families like Felidae and Cancidae. Class The group of related orders is called class.  For example; order Primata comprising monkey, gorilla and gibbon is placed in class Mammalia along with order Carnivora that includes animals like tiger, cat and dog. Presence of mammary glands is one of the common characters in these orders. Phylum In the Animal Kingdom; group of closely related classes is called phylum. In the Plant Kingdom; group of closely related classes is called Division. For example; fishes, amphibians, reptiles, birds along with mammals belong to Phylum Chordata. The common features for Chordata are; presence of notochord and dorsal hollow neural system. Kingdom The group of all related phyla is called Kingdom. For example; all heterotrophs; which do not have cell wall and are eukaryotic are kept under the Animal Kingdom. All autotrophs which are eukaryotic are kept under Plant Kingdom.
Test Your Knowledge 1. Which is the lowest taxa in the classification of living beings?  2. Why do fish and mammals belong to the same Phylum?

BIOLOGICAL CLASSIFICATION Five Kingdom Classification as proposed by R.H. Whittaker (1969): 1. Monera, 2. Protista, 3. Fungi, 4. Plantae and 5. Animalia. The main criteria for classification used by Whittaker: 1. Cell Structure 2. Thallus Organization 3. Mode of Nutrition 4. Reproduction and 5. Phylogenetic Relationships KINGDOM MONERA Bacteria are grouped under four categories based on their shape: 1. The spherical Coccus (pl.: cocci), 2. The rod-shaped Bacillus (pl.: bacilli), 3. The comma-shaped Vibrium (pl.: vibrio) and 4. The spiral Spirillum (pl.: spirilla) Though the bacterial structure is very simple, they are very complex in behaviour. Compared to many other organisms, bacteria as a group show the most extensive metabolic diversity. Some of the bacteria are autotrophic, i.e., they synthesise their own food from inorganic substrates. They may be photosynthetic autotrophic or chemosynthetic autotrophic. The vast majority of bacteria are heterotrophs, i.e., they do not synthesise their own food but depend on other organisms or on dead organic matter for food. Archaebacteria These bacteria are special since they live in some of the most harsh habitats and named according to their habitats as follows: 1. Halophiles live in extreme salty areas, 2. Thermoacidophiles live in hot springs and 3. Methanogens live in marshy areas Archaebacteria differ from other bacteria in having a different cell wall structure and this feature is responsible for their survival in extreme conditions. Methanogens are present in the guts of several ruminant animals such as cows and buffaloes and they are responsible for the production of methane (biogas) from the dung of these animals. Eubacteria There are thousands of different eubacteria or ‘true bacteria’. They are characterised by the presence of a rigid cell wall, and if motile, a flagellum. The cyanobacteria (also referred to as blue-green algae) have chlorophyll a similar to green plants and are photosynthetic autotrophs. The cyanobacteria are unicellular, colonial or filamentous, marine or terrestrial algae. The colonies are generally surrounded by gelatinous sheath. They often form blooms in polluted water bodies. Some of these organisms can fix atmospheric nitrogen in specialised cells called heterocysts, e.g., Nostoc and Anabaena. Chemosynthetic autotrophic bacteria oxidise various inorganic substances such as nitrates, nitrites and ammonia and use the released energy for their ATP production. They play a great role in recycling nutrients like nitrogen, phosphorous, iron and sulphur. Heterotrophic bacteria are the most abundant in nature. The majority are important decomposers. Many of them have a significant impact on human affairs. They are helpful in making curd from milk, production of antibiotics, fixing nitrogen in legume roots, etc. Some are pathogens causing damage to human beings, crops, farm animals and pets. Cholera, typhoid, tetanus, citrus canker are well known diseases caused by different bacteria. Bacteria reproduce mainly by fission. Sometimes, under unfavourable conditions, they produce spores. They also reproduce by a sort of sexual reproduction by adopting a primitive type of DNA transfer from one bacterium to the other. The Mycoplasmas are organisms that completely lack a cell wall. They are the smallest living cells known and can survive without oxygen. Many mycoplasma are pathogenic in animals and plants. KINGDOM PROTISTA All single-celled eukaryotes are placed under Protista, but the boundaries of this kingdom are not well defined. Being eukaryotes, the protistan cell body contains a well defined nucleus and other membrane-bound organelles. Some have flagella or cilia. Protists reproduce asexually and sexually by a process involving cell fusion and zygote formation. Chrysophytes This group includes diatoms and golden algae (desmids). They are found in fresh water as well as in marine environments. They are microscopic and float passively in water currents (plankton). Most of them are photosynthetic. In diatoms the cell walls form two thin overlapping shells, which fit together as in a soap box. The walls are embedded with silica and thus the walls are indestructible. Thus, diatoms have left behind large amount of cell wall deposits in their habitat; this accumulation over billions of years is referred to as ‘diatomaceous earth’. Being gritty this soil is used in polishing, filtration of oils and syrups. Diatoms are the chief ‘producers’ in the oceans. Dianoflagellates These organisms are mostly marine and photosynthetic. They appear yellow, green, brown, blue or red depending on the main pigments present in their cells. The cell wall has stiff cellulose plates on the outer surface. Most of them have two flagella; one lies longitudinally and the other transversely in a furrow between the wall plates. Euglenoids Majority of them are fresh water organisms found in stagnant water. Instead of a cell wall, they have a protein rich layer called pellicle which makes their body flexible. They have two flagella, a short and a long one. Though they are photosynthetic in the presence of sunlight, when deprived of sunlight they behave like heterotrophs by predating on other smaller organisms. Example: Euglena. Slime Moulds Slime moulds are saprophytic protists. The body moves along decaying twigs and leaves engulfing organic material. Under suitable conditions, they form an aggregation called plasmodium which may grow and spread over several feet. During unfavourable conditions, the plasmodium differentiates and forms fruiting bodies bearing spores at their tips. The spores possess true walls. They are extremely resistant and survive for many years, even under adverse conditions. The spores are dispersed by air currents. Protozoans All protozoans are heterotrophs and live as predators or parasites. There are four major groups of protozoans. 1. Amoeboid protozoans: These organisms live in fresh water, sea water or moist soil. They move and capture their prey by putting out pseudopodia (false feet) as in Amoeba. Marine forms have silica shells on their surface. Some of them such as Entamoeba are parasites. 2. Flagellated protozoans: The members of this group are either free-living or parasitic. They have flagella. The parasitic forms cause diaseases such as sleeping sickness. Example: Trypanosoma. 3. Ciliated protozoans: These are aquatic, actively moving organisms because of the presence of thousands of cilia. They have a cavity (gullet) that opens to the outside of the cell surface. The coordinated movement of rows of cilia causes the water laden with food to be steered into the gullet. Example: Paramoecium 4. Sporozoans: This includes diverse organisms that have an infectious spore-like stage in their life cycle. The most notorious is Plasmodium (malarial parasite) which causes malaria which has a staggering effect on human population. KINGDOM FUNGI With the exception of yeasts which are unicellular, fungi are filamentous. Their bodies consist of long, slender thread-like structures called hyphae. The network of hyphae is known as mycelium. Some hyphae are continuous tubes filled with multinucleated cytoplasm – these are called coenocytic hyphae. Others have septae or cross walls in their hyphae. The cell walls of fungi are composed of chitin and polysaccharides. Most fungi are heterotrophic and absorb soluble organic matter from dead substrates and hence are called saprophytes. Those that depend on living plants and animals are called parasites. They can also live as symbionts – in association with algae as lichens and with roots of higher plants as mycorrhiza. Reproduction in Fungi: Reproduction in fungi can take place by vegetative means – fragmentation, fission and budding. Asexual reproduction is by spores called conidia or sporangiospores or zoospores, and sexual reproduction is by oospores, ascospores and basidiospores. The various spores are produced in distinct structures called fruiting bodies. The sexual cycle involves the following three steps: (i) Fusion of protoplasms between two motile or non-motile gametes called plasmogamy. (ii) Fusion of two nuclei called karyogamy. (iii) Meiosis in zygote resulting in haploid spores. When a fungus reproduces sexually, two haploid hyphae of compatible mating types come together and fuse. In some fungi the fusion of two haploid cells immediately results in diploid cells (2n). However, in other fungi (ascomycetes and basidiomycetes), an intervening dikaryotic stage (n + n i.e. two nuclei per cell) occurs; such a condition is called a dikaryon and the phase is called dikaryophase of fungus. Later, the parental nuclei fuse and the cells become diploid. The fungi form fruiting bodies in which reduction division occurs, leading to formation of haploid spores. The morphology of the mycelium, mode of spore formation and fruiting bodies form the basis for the division of the kingdom into various classes. Phycomycetes Members of phycomycetes are found in aquatic habitats and on decaying wood in moist and damp places or as obligate parasites on plants. Mycelium: The mycelium is aseptate and coenocytic. Reproduction: Asexual reproduction takes place by zoospores (motile) or by aplanospores (non-motile). These spores are endogeneously produced in sporangium. Zygospores are formed by fusion of two gametes. These gametes are similar in morphology (isogamous) or dissimilar (anisogamous or oogamous). Some common examples are Mucor, Rhizopus (the bread mould mentioned earlier) and Albugo (the parasitic fungi on mustard). Ascomycetes Commonly known as sac-fungi, the ascomycetes are unicellular, e.g., yeast (Sacharomyces) or multicellular, e.g., Penicillium. They are saprophytic, decomposers, parasitic or coprophilous (growing on dung). Mycelium: Mycelium is branched and septate. Reproduction: The asexual spores are conidia produced exogenously on the special mycelium called conidiophores. Conidia on germination produce mycelium. Sexual spores are called ascospores which are produced endogenously in sac like asci (singular ascus). These asci are arranged in different types of fruiting bodies called ascocarps. Some examples are Aspergillu, Claviceps and Neurospora. Neurospora is used extensively in biochemical and genetic work. Many members like morels and buffles are edible and are considered delicacies. Basidiomycetes Commonly known forms of basidiomycetes are mushrooms, bracket fungi or puffballs. They grow in soil, on logs and tree stumps and in living plant bodies as parasites, e.g., rusts and smuts. Mycelium: The mycelium is branched and septate. Reproduction: The asexual spores are generally not found, but vegetative reproduction by fragmentation is common. The sex organs are absent, but plasmogamy is brought about by fusion of two vegetative or somatic cells of different strains or genotypes. The resultant structure is dikaryotic which ultimately gives rise to basidium. Karyogamy and meiosis take place in the basidium producing four basidiospores. The basidiospores are exogenously produced on the basidium (pl.: basidia). The basidia are arranged in fruiting bodies called basidiocarps. Some common members are Agaricus (mushroom), Ustilago (smut) and Puccinia (rust fungus). Deuteromycetes Commonly known as imperfect fungi, because only the asexual or vegetative phases of these fungi are known. When the sexual forms of these fungi were discovered they were moved into classes they rightly belong to. It is also possible that the asexual and vegetative stage have been given one name (and placed under deuteromycetes) and the sexual stage another (and placed under another class). Later when the linkages were established, the fungi were correctly identified and moved out of deuteromycetes. Once perfect (sexual) stages of members of dueteromycetes were discovered they were often moved to ascomycetes and basidiomycetes. The deuteromycetes reproduce only by asexual spores known as conidia. The mycelium is septate and branched. Some members are saprophytes or parasites while a large number of them are decomposers of litter and help in mineral cycling. Some examples are Alternaria, Colletotrichum and Trichoderma. KINGDOM PLANTAE Kingdom Plantae includes all eukaryotic chlorophyll-containing organisms commonly called plants. A few members are partially heterotrophic such as the insectivorous plants or parasites. Bladderwort and Venus fly trap are examples of insectivorous plants and Cuscuta is a parasite. The plant cells have an eukaryotic structure with prominent chloroplasts and cell wall mainly made of cellulose. Plantae includes algae, bryophytes, pteridophytes, gymnosperms and angiosperms. Life cycle of plants has two distinct phases – the diploid sporophytic and the haploid gametophytic – that alternate with each other. The lengths of the haploid and diploid phases, and whether these phases are free– living or dependent on others, vary among different groups in plants. This phenomenon is called alternation of generation. KINGDOM ANIMALIA This kingdom is characterised by heterotrophic eukaryotic organisms that are multicellular and their cells lack cell walls. They directly or indirectly depend on plants for food. They digest their food in an internal cavity and store food reserves as glycogen or fat. Their mode of nutrition is holozoic – by ingestion of food. They follow a definite growth pattern and grow into adults that have a definite shape and size. Higher forms show elaborate sensory and neuromotor mechanism. Most of them are capable of locomotion. The sexual reproduction is by copulation of male and female followed by embryological development. VIRUSES, VIROIDS AND LICHENS Viruses did not find a place in classification since they are not truly ‘living’, if we understand living as those organisms that have a cell structure. The viruses are non-cellular organisms that are characterized by having an inert crystalline structure outside the living cell. Once they infect a cell they take over the machinery of the host cell to replicate themselves, killing the host. The name virus that means venom or poisonous fluid was given by Pasteur. D.J. Ivanowsky (1892) recognised certain microbes as causal organism of the mosaic disease of tobacco. These were found to be smaller than bacteria because they passed through bacteria-proof filters. M.W. Beijerinek (1898) demonstrated that the extract of the infected plants of tobacco could cause infection in healthy plants and called the fluid as Contagium vivum fluidum (infectious living fluid). W.M. Stanley (1935) showed that viruses could be crystallised and crystals consist largely of proteins. They are inert outside their specific host cell. Viruses are obligate parasites. In addition to proteins viruses also contain genetic material, that could be either RNA or DNA. No virus contains both RNA and DNA. A virus is a nucleoprotein and the genetic material is infectious. In general, viruses that infect plants have single stranded RNA and viruses that infect animals have either single or double stranded RNA or double stranded DNA. Bacterial viruses or bacteriophages (viruses that infect the bacteria) are usually double stranded DNA viruses. The protein coat called capsid made of small subunits called capsomeres, protects the nucleic acid. These capsomeres are arranged in helical or polyhedral geometric forms. Viruses cause diseases like mumps, small pox, herpes and influenza. AIDS in humans is also caused by a virus. In plants, the symptoms can be mosaic formation, leaf rolling and curling, yellowing and vein clearing, dwarfing and stunted growth.

PLANT KINGDOM At present phylogenetic classification systems based on evolutionary relationships between the various organisms are acceptable. This assumes that organisms belonging to the same taxa have a common ancestor. We now use information from many other sources too to help resolve difficulties in classification. These become more important when there is no supporting fossil evidence. Numerical Taxonomy is based on all observable characteristics. Number and codes are assigned to all the characters and the data are then processed. In this way each character is given equal importance and at the same time hundreds of characters can be considered. Cytotaxonomy that is based on cytological information like chromosome number, structure, behaviour and chemotaxonomy that uses the chemical constituents of the plant to resolve confusions, are also used by taxonomists these days. ALGAE Habit & Habitat: Algae are chlorophyll-bearing, simple, thalloid, autotrophic and largely aquatic (both fresh water and marine) organisms. They occur in a variety of other habitats: moist stones, soils and wood. Some of them also occur in association with fungi (lichen) and animals (e.g., on sloth bear). Size: The size ranges from the microscopic unicellular forms like Chlamydomonas, to colonial forms like Volvox and to the filamentous forms like Ulothrix and Spirogyra. A few of the marine forms, such as kelps, form massive plant bodies. Reproduction: The algae reproduce by vegetative, asexual and sexual methods. Vegetative Reproduction: Vegetative reproduction is by fragmentation. Each fragment develops into a thallus. Asexual Reproduction: Asexual reproduction is by the production of different types of spores, the most common being the zoospores. They are flagellated (motile) and on germination gives rise to new plants. Sexual reproduction: Sexual reproduction takes place through fusion of two gametes. The fusion of gametes can be of following types in algae: Isogamous Fusion: These gametes can be flagellated and similar in size (as in Chlamydomonas) or non-flagellated (non-motile) but similar in size (as in Spirogyra). Such reproduction is called isogamous. Anisogamous Fusion: Fusion of two gametes dissimilar in size, as in some species of Chlamydomonas is termed as anisogamous. Oogamous Fusion: Fusion between one large, non-motile (static) female gamete and a smaller, motile male gamete is termed oogamous, e.g., Volvox, Fucus. Economic Importance of Algae: Algae are useful to man in a variety of ways. At least a half of the total carbon dioxide fixation on earth is carried out by algae through photosynthesis. Being photosynthetic they increase the level of dissolved oxygen in their immediate environment. They are of paramount importance as primary producers of energy-rich compounds which form the basis of the food cycles of all aquatic animals. Many species of Porphyra, Laminaria and Sargassum are among the 70 species of marine algae used as food. Certain marine brown and red algae produce large amounts of hydrocolloids (water holding substances), e.g., algin (brown algae) and carrageen (red algae) are used commercially. Agar, one of the commercial products obtained from Gelidium and Gracilaria are used to grow microbes and in preparations of ice-creams and jellies. Chlorella and Spirullina are unicellular algae, rich in proteins and are used as food supplements even by space travellers. The algae are divided into three main classes: Chlorophyceae, Phaeophyceae and Rhodophyceae. Chlorophyceae Characteristics: The members of chlorophyceae are commonly called green algae. The plant body may be unicellular, colonial or filamentous. They are usually grass green due to the dominance of pigments chlorophyll a and b. The pigments are localised in definite chloroplasts. The chloroplasts may be discoid, plate-like, reticulate, cup-shaped, spiral or ribbon-shaped in different species. Most of the members have one or more storage bodies called pyrenoids located in the chloroplasts. Pyrenoids contain protein besides starch. Some algae may store food in the form of oil droplets. Green algae usually have a rigid cell wall made of an inner layer of cellulose and an outer layer of pectose. Reproduction: Vegetative reproduction usually takes place by fragmentation or by formation of different types of spores. Asexual reproduction is by flagellated zoospores produced in zoosporangia. The sexual reproduction shows considerable variation in the type and formation of sex cells and it may be isogamous, anisogamous or oogamous. Common Examples: Chlamydomonas, Volvox, Ulothrix, Spirogyra and Chara Phaeophyceae Characteristics: The members of phaeophyceae or brown algae are found primarily in marine habitats. They show great variation in size and form. They range from simple branched, filamentous forms (Ectocarpus) to profusely branched forms as represented by kelps, which may reach a height of 100 metres. They possess chlorophyll a, c, carotenoids and xanthophylls. They vary in colour from olive green to various shades of brown depending upon the amount of the xanthophyll pigment, fucoxanthin present in them. Food is stored as complex carbohydrates, which may be in the form of laminarin or mannitol. The vegetative cells have a cellulosic wall usually covered on the outside by a gelatinous coating of algin. The protoplast contains, in addition to plastids, a centrally located vacuole and nucleus. The plant body is usually attached to the substratum by a holdfast, and has a stalk, the stipe and leaf like photosynthetic organ – the frond. Vegetative reproduction takes place by fragmentation. Reproduction: Asexual reproduction in most brown algae is by biflagellate zoospores that are pear-shaped and have two unequal laterally attached flagella. Sexual reproduction may be isogamous, anisogamous or oogamous. Union of gametes may take place in water or within the oogonium (oogamous species). The gametes are pyriform (pear-shaped) and bear two laterally attached flagella. Common Examples: Ectocarpus, Dictyota, Laminaria, Sargassum and Fucus Rhodophyceae Characteristics: Rhodophyta are commonly called red algae because of the predominance of the red pigment, r-phycoerythrin in their body. Majority of the red algae are marine with greater concentrations found in the warmer areas. They occur in both well-lighted regions close to the surface of water and also at great depths in oceans where relatively little light penetrates. The red thalli of most of the red algae are multicellular. Some of them have complex body organisation. The food is stored as floridean starch which is very similar to amylopectin and glycogen in structure. Reprodcution: The red algae usually reproduce vegetatively by fragmentation. They reproduce asexually by non-motile spores and sexually by non-motile gametes. Sexual reproduction is oogamous and accompanied by complex post fertilisation developments. Common Examples: Polysiphonia, Porphyra, Gracilaria and Gelidium. BRYOPHYTES Habits & Habitats: Bryophytes include the various mosses and liverworts that are found commonly growing in moist shaded areas in the hills. Bryophytes are also called amphibians of the plant kingdom because these plants can live in soil but are dependent on water for sexual reproduction. They usually occur in damp, humid and shaded localities. Characteristics: They play an important role in plant succession on bare rocks/soil. The plant body of bryophytes is more differentiated than that of algae. It is thallus-like and prostrate or erect, and attached to the substratum by unicellular or multicellular rhizoids. They lack true roots, stem or leaves. They may possess root-like, leaf-like or stem-like structures. Reproduction: The main plant body of the bryophyte is haploid. It produces gametes, hence is called a gametophyte. The sex organs in bryophytes are multicellular. The male sex organ is called antheridium. They produce biflagellate antherozoids. The female sex organ called archegonium is flask-shaped and produces a single egg. The antherozoids are released into water where they come in contact with archegonium. An antherozoid fuses with the egg to produce the zygote. Zygotes do not undergo reduction division immediately. They produce a multicellular body called a sporophyte. The sporophyte is not free-living but attached to the photosynthetic gametophyte and derives nourishment from it. Some cells of the sporophyte undergo reduction division (meiosis) to produce haploid spores. These spores germinate to produce gametophyte. Economic Importance of Bryophytes: Bryophytes in general are of little economic importance but some mosses provide food for herbaceous mammals, birds and other animals. Species of Sphagnum, a moss, provide peat that have long been used as fuel, and because of their capacity to hold water as packing material for trans-shipment of living material. Mosses along with lichens are the first organisms to colonise rocks and hence, are of great ecological importance. They decompose rocks making the substrate suitable for the growth of higher plants. Since mosses form dense mats on the soil, they reduce the impact of falling rain and prevent soil erosion. The bryophytes are divided into following classes: 1. liverworts and 2. mosses. Liverworts Characteristics: The liverworts grow usually in moist, shady habitats such as banks of streams, marshy ground, damp soil, bark of trees and deep in the woods. The plant body of a liverwort is thalloid, e.g., Marchantia. The thallus is dorsiventral and closely appressed to the substrate. The leafy members have tiny leaf-like appendages in two rows on the stem-like structures. Reproduction: Asexual reproduction in liverworts takes place by fragmentation of thalli, or by the formation of specialised structures called gemmae (sing. gemma). Gemmae are green, multicellular, asexual buds, which develop in small receptacles called gemma cups located on the thalli. The gemmae become detached from the parent body and germinate to form new individuals. During sexual reproduction, male and female sex organs are produced either on the same or on different thalli. The sporophyte is differentiated into a foot, seta and capsule. After meiosis, spores are produced within the capsule. These spores germinate to form free-living gametophytes. Mosses Characteristics: The predominant stage of the life cycle of a moss is the gametophyte which consists of two stages. The first stage is the protonema stage, which develops directly from a spore. It is a creeping, green, branched and frequently filamentous stage. The second stage is the leafy stage, which develops from the secondary protonema as a lateral bud. They consist of upright, slender axes bearing spirally arranged leaves. They are attached to the soil through multicellular and branched rhizoids. This stage bears the sex organs. Reproduction: Vegetative reproduction in mosses is by fragmentation and budding in the secondary protonema. In sexual reproduction, the sex organs antheridia and archegonia are produced at the apex of the leafy shoots. After fertilisation, the zygote develops into a sporophyte, consisting of a foot, seta and capsule. The sporophyte in mosses is more elaborate than that in liverworts. The capsule contains spores. Spores are formed after meiosis. The mosses have an elaborate mechanism of spore dispersal. Common Examples: Funaria, Polytrichum and Sphagnum PTERIDOPHYTES Characteristics: The pteridophytes are found in cool, damp, shady places though some may flourish well in sandy-soil conditions. In pteridophytes, the main plant body is a sporophyte which is differentiated into true root, stem and leaves. These organs possess well-differentiated vascular tissues. The leaves in pteridophyta are small (microphylls) as in Selaginella or large (macrophylls) as in ferns. The sporophytes bear sporangia that are subtended by leaf-like appendages called sporophylls. In some cases sporophylls may form distinct compact structures called strobili or cones (Selaginella, Equisetum). Reproduction: The sporangia produce spores by meiosis in spore mother cells. The spores germinate to give rise to inconspicuous, small but multicellular, free-living, mostly photosynthetic thalloid gametophytes called prothallus. These gametophytes require cool, damp, shady places to grow. Because of this specific restricted requirement and the need for water for fertilisation, the spread of living pteridophytes is limited and restricted to narrow geographical regions. The gametophytes bear male and female sex organs called antheridia and archegonia, respectively. Water is required for transfer of antherozoids – the male gametes released from the antheridia, to the mouth of archegonium. Fusion of male gamete with the egg present in the archegonium result in the formation of zygote. Zygote thereafter produces a multicellular well-differentiated sporophyte which is the dominant phase of the pteridophytes. In majority of the pteridophytes all the spores are of similar kinds; such plants are called homosporous. Genera like Selaginella and Salvinia which produce two kinds of spores, macro (large) and micro (small) spores, are known as heterosporous. The megaspores and microspores germinate and give rise to female and male gametophytes, respectively. The female gametophytes in these plants are retained on the parent sporophytes for variable periods. The development of the zygotes into young embryos take place within the female gametophytes. This event is a precursor to the seed habit considered an important step in evolution. The pteridophytes are divided into four classes: 1. Psilopsida(Psilotum); 2. Lycopsida (Selaginella, Lycopodium), 3. Sphenopsida (Equisetum) and 4. Pteropsida (Dryopteris, Pteris, Adiantum). GYMNOSPERMS Characteristics: The gymnosperms (gymnos : naked, sperma : seeds) are plants in which the ovules are not enclosed by any ovary wall and remain exposed, both before and after fertilisation. The seeds that develop post-fertilisation, are not covered, i.e., are naked. Gymnosperms include medium-sized trees or tall trees and shrubs. One of the gymnosperms, the giant redwood tree Sequoia is one of the tallest tree species. The roots are generally tap roots. Roots in some genera have fungal association in the form of mycorrhiza (Pinus), while in some others (Cycas) small specialized roots called coralloid roots are associated with N2- fixing cyanobacteria. The stems are unbranched (Cycas) or branched (Pinus, Cedrus). The leaves may be simple or compound. In Cycas the pinnate leaves persist for a few years. The leaves in gymnosperms are well-adapted to withstand extremes of temperature, humidity and wind. In conifers, the needle-like leaves reduce the surface area. Their thick cuticle and sunken stomata also help to reduce water loss. Reproduction: The gymnosperms are heterosporous; they produce haploid microspores and megaspores. The two kinds of spores are produced within sporangia that are borne on sporophylls which are arranged spirally along an axis to form lax or compact strobili or cones. The strobili bearing microsporophylls and microsporangia are called microsporangiate or male strobili. The microspores develop into a male gametophytic generation which is highly reduced and is confined to only a limited number of cells. This reduced gametophyte is called a pollen grain. The development of pollen grains takes place within the microsporangia. The cones bearing megasporophylls with ovules or megasporangia are called macrosporangiate or female strobili. The male or female cones or strobili may be borne on the same tree (Pinus) or on different trees (Cycas). The megaspore mother cell is differentiated from one of the cells of the nucellus. The nucellus is protected by envelopes and the composite structure is called an ovule. The ovules are borne on megasporophylls which may be clustered to form the female cones. The megaspore mother cell divides meiotically to form four megaspores. One of the megaspores enclosed within the megasporangium (nucellus) develops into a multicellular female gametophyte that bears two or more archegonia or female sex organs. The multicellular female gametophyte is also retained within megasporangium. Fertilization: Unlike bryophytes and pteridophytes, in gymnosperms the male and the female gametophytes do not have an independent free-living existence. They remain within the sporangia retained on the sporophytes. The pollen grain is released from the microsporangium. They are carried in air currents and come in contact with the opening of the ovules borne on megasporophylls. The pollen tube carrying the male gametes grows towards archegonia in the ovules and discharge their contents near the mouth of the archegonia. Following fertilisation, zygote develops into an embryo and the ovules into seeds. These seeds are not covered. ANGIOSPERMS Characters: In the angiosperms or flowering plants, the pollen grains and ovules are developed in specialized structures called flowers. In angiosperms, the seeds are enclosed by fruits. The angiosperms are an exceptionally large group of plants occurring in wide range of habitats. They range in size from tiny, almost microscopic Wolfia to tall trees of Eucalyptus (over 100 metres). They provide us with food, fodder, fuel, medicines and several other commercially important products. Angiosperms are divided into two classes: 1. Dicotyledons and 2. Monocotyledons The dicotyledons are characterised by having two cotyledons in their seeds while the monocolyledons have only one. Reproduction: Male Sex Organ: The male sex organ in a flower is the stamen. Each stamen consists of a slender filament with an anther at the tip. The anthers, following meiosis, produce pollen grains. Female Sex Organs: The female sex organs in a flower is the pistil or the carpel. Pistil consists of an ovary enclosing one to many ovules. Within ovules are present highly reduced female gametophytes termed embryosacs. The embryo-sac formation is preceded by meiosis. Hence, each of the cells of an embryo-sac is haploid. Each embryo-sac has a three-celled egg apparatus – one egg cell and two synergids, three antipodal cells and two polar nuclei. The polar nuclei eventually fuse to produce a diploid secondary nucleus. Pollination: Pollen grains, after dispersal from the anthers, are carried by wind or various other agencies to the stigma of a pistil. This is termed as pollination. Fertilization: The pollen grains germinate on the stigma and the resulting pollen tubes grow through the tissues of stigma and style and reach the ovule. The pollen tubes enter the embryo-sac where two male gametes are discharged. One of the male gametes fuses with the egg cell to form a zygote (syngamy). The other male gamete fuses with the diploid secondary nucleus to produce the triploid primary endosperm nucleus (PEN). Because of the involvement of two fusions, this event is termed as double fertilisation, an event unique to angiosperms. Embryo: The zygote develops into an embryo (with one or two cotyledons) and the PEN develops into endosperm which provides nourishment to the developing embryo. The synergids and antipodals degenerate after fertilisation. During these events the ovules develop into seeds and the ovaries develop into fruit. PLANT LIFE CYCLES AND ALTERNATION OF GENERATIONS In plants, both haploid and diploid cells can divide by mitosis. This ability leads to the formation of different plant bodies - haploid and diploid. The haploid plant body produces gametes by mitosis. This plant body represents a gametophyte. Following fertilisation the zygote also divides by mitosis to produce a diploid sporophytic plant body. Haploid spores are produced by this plant body by meiosis. These in turn, divide by mitosis to form a haploid plant body once again. Thus, during the life cycle of any sexually reproducing plant, there is an alternation of generations between gamete producing haploid gametophyte and spore producing diploid sporophyte. However, different plant groups, as well as individuals representing them, differ in the following patterns: 1. Sporophytic generation is represented only by the one-celled zygote. There are no free-living sporophytes. Meiosis in the zygote results in the formation of haploid spores. The haploid spores divide mitotically and form the gametophyte. The dominant, photosynthetic phase in such plants is the free-living gametophyte. This kind of life cycle is termed as haplontic. Many algae such as Volvox, Spirogyra and some species of Chlamydomomas represent this pattern. 2. On the other extreme, is the type wherein the diploid sporophyte is the dominant, photosynthetic, independent phase of the plant. The gametophytic phase is represented by the single to few-celled haploid gametophyte. This kind of lifecycle is termed as diplontic. All seed-bearing plants i.e. gymnosperms and angiosperms, follow this pattern. 3. Bryophytes and pteridophytes, interestingly, exhibit an intermediate condition (Haplo-diplontic); both phases are multicellular and often free-living. However, they differ in their dominant phases.

Animal Kingdom Basis Of Classification Levels of Organisation 1. Cellular Level: In unicellular animals and some multicellular animals, the cell is responsible for all the metabolic activities in the animal body. This type of organization of function is termed as cellular level of organization. 2. Tissue Level: In certain animals cells performing the same function are arranged in a group. Example: Coelenterates. 3. Organ Level: Some animals form specialized organs for specific functions. Example: Platyhelminthes. 4. Organ System Level: In higher and complex animals various organs group to form a complex organ system to perform specific function. Example: Molluscs, Chordata. Symmetry Animals can be categorised on the basis of their symmetry. Sponges are mostly asymmetrical, i.e., any plane that passes through the centre does not divide them into equal halves. Radial Symmetry: When any plane passing through the central axis of the body divides the organism into two identical halves, it is called radial symmetry. Coelenterates, ctenophores and echinoderms have this kind of body plan. Bilateral Symmetry: Animals like annelids, arthropods, etc., where the body can be divided into identical left and right halves in only one plane, exhibit bilateral symmetry. Diploblastic and Triploblastic Organisation Animals in which the cells are arranged in two embryonic layers, an external ectoderm and an internal endoderm, are called diploblastic animals, e.g., coelenterates. An undifferentiated layer, mesoglea, is present in between the ectoderm and the endoderm. Those animals in which the developing embryo has a third germinal layer, mesoderm, in between the ectoderm and endoderm, are called triploblastic animals (platyhelminthes to chordates). Coelom Coelomates: Presence or absence of a cavity between the body wall and the gut wall is very important in classification. The body cavity, which is lined by mesoderm is called coelom. Animals possessing coelom are called coelomates, e.g., annelids, molluscs, arthropods, echinoderms, hemichordates and chordates. Pseudoceolomates: In some animals, the body cavity is not lined by mesoderm, instead, the mesoderm is present as scattered pouches in between the ectoderm and endoderm. Such a body cavity is called pseudocoelom and the animals possessing them are called pseudocoelomates, e.g., aschelminthes. Acoelomates: The animals in which the body cavity is absent are called acoelomates, e.g., platyhelminthes. Segmentation In some animals, the body is externally and internally divided into segments with a serial repetition of at least some organs. For example, in earthworm, the body shows this pattern called metameric segmentation and the phenomenon is known as metamerism. Notochord Notochord is a mesodermally derived rod-like structure formed on the dorsal side during embryonic development in some animals. Animals with notochord are called chordates and those animals which do not form this structure are called non-chordates, e.g., porifera to echinoderms. CLASSIFICATION OF ANIMALS Phylum – Porifera Members of this phylum are commonly known as sponges. They are generally marine and mostly asymmetrical animals. These are primitive multicellular animals and have cellular level of organisation. Water Transport System in Sponges: Sponges have a water transport or canal system. Water enters through minute pores (ostia) in the body wall into a central cavity, spongocoel, from where it goes out through the osculum. This pathway of water transport is helpful in food gathering, respiratory exchange and removal of waste. Choanocytes or collar cells line the spongocoel and the canals. Digestion is intracellular. The body is supported by a skeleton made up of spicules or sponging fibres. Reproduction: Sponges are hermaphrodite animals. Sexes are not separate, i.e., eggs and sperms are produced by the same individual. Sponges reproduce asexually by fragmentation and sexually by formation of gametes. Fertilisation is internal and development is indirect having a larval stage which is morphologically distinct from the adult. Phylum – Coelenterata (Cnidaria) They are aquatic, mostly marine, sessile or free-swimming, radially symmetrical animals. The name cnidaria is derived from the nidoblasts or cnidocytes (which contain the stinging capsules or nematocytes) present on the tentacles and the body. Cnidoblasts are used for anchorage, defense and for the capture of prey. Cnidarians exhibit tissue level of organisation and are diploblastic. They have a central gastro-vascular cavity with a single opening, hypostome. Digestion is extracellular and intracellular. Some of the cnidarians, e.g., corals have a skeleton composed of calcium carbonate.   :  Cnidarians exhibit two basic body forms called polyp and medusa. Polyp is a sessile and cylindrical form like Hydra, Adamsia, etc. whereas, medussa is umbrella-shaped and free-swimming like Aurelia or jelly fish. Those cnidarians which exist in both forms exhibit alternation of generation (Metagenesis), i.e., polyps produce medusae asexually and medusae form the polyps sexually (e.g., Obelia). Examples: Physalia (Portuguese man-of-war), Adamsia (Sea anemone), Pennatula (Sea-pen), Gorgonia (Sea-fan) and Meandrina (Brain coral). Phylum – Ctenophora Ctenophores, commonly known as sea walnuts or comb jellies are exclusively marine, radially symmetrical, diploblastic organisms with tissue level of organisation. The body bears eight external rows of ciliated comb plates, which help in locomotion. Digestion is both extracellular and intracellular. Bioluminescence (the property of a living organism to emit light) is well-marked in ctenophores. Sexes are not separate. Reproduction takes place only by sexual means. Fertilisation is external with indirect development. Examples: Pleurobrachia and Ctenoplana. Phylum – Platyhelminthes They have dorso-ventrally flattened body, hence are called flatworms. These are mostly endoparasites found in animals including human beings. Flatworms are bilaterally symmetrical, triploblastic and acoelomate animals with organ level of organisation. Hooks and suckers are present in the parasitic forms. Some of them absorb nutrients from the host directly through their body surface. Specialised cells called flame cells help in osmoregulation and excretion. Sexes are not separate. Fertilisation is internal and development is through many larval stages. Some members like Planaria possess high regeneration capacity. Examples: Taenia (Tapeworm), Fasciola (Liver fluke). Phylum – Aschelminthes The body of the aschelminthes is circular in cross-section, hence, the name roundworms. They may be freeliving, aquatic and terrestrial or parasitic in plants and animals. Roundworms have organ-system level of body organisation. They are bilaterally symmetrical, triploblastic and pseudocoelomate animals. Digestive System: Alimentary canal is complete with a well developed muscular pharynx. An excretory tube removes body wastes from the body cavity through the excretory pore. Sexes are separate (dioecious), i.e., males and females are distinct. Reproduction: Often females are longer than males. Fertilisation is internal and development may be direct (the young ones resemble the adult) or indirect. Examples: Ascaris (Round Worm), Wuchereria (Filaria worm), Ancylostoma (Hookworm). Phylum – Annelida They may be aquatic (marine and fresh water) or terrestrial; free-living, and sometimes parasitic. They exhibit organ-system level of body organisation and bilateral symmetry. They are triploblastic, metamerically segmented and coelomate animals. Their body surface is distinctly marked out into segments or metameres (Latin, annulus : little ring) and, hence, the phylum name Annelida. They possess longitudinal and circular muscles which help in locomotion. Aquatic annelids like Nereis possess lateral appendages, parapodia, which help in swimming. A closed circulatory system is present. Nephridia (sing. nephridium) help in osmoregulation and excretion. Neural system consists of paired ganglia (sing. ganglion) connected by lateral nerves to a double ventral nerve cord. Nereis, an aquatic form, is dioecious, but earthworms and leeches are monoecious. Reproduction is sexual. Examples: Nereis, Pheretima (Earthworm) and Hirudinaria (Blood sucking leech). Male Female Phylum – Arthropoda This is the largest phylum of Animalia which includes insects. Over two-thirds of all named species on earth are arthropods. They have organ-system level of organisation. They are bilaterally symmetrical, triploblastic, segmented and coelomate animals. Structure: The body of arthropods is covered by chitinous exoskeleton. The body consists of head, thorax and abdomen. They have jointed appendages (arthros-joint, poda-appendages). Respiratory organs are gills, book gills, book lungs or tracheal system. Circulatory system is of open type. Sensory organs like antennae, eyes (compound and simple), statocysts or balance organs are present. Excretion takes place through malpighian tubules. They are mostly dioecious. Fertilisation is usually internal. They are mostly oviparous. Development may be direct or indirect. Examples: Economically important insects – Apis (Honey bee), Bombyx (Silkworm), Laccifer (Lac insect) Vectors – Anopheles, Culex and Aedes (Mosquitoes) Gregarious pest – Locusta (Locust) Living fossil – Limulus (King crab). Phylum – Mollusca This is the second largest animal phylum. Molluscs are terrestrial or aquatic (marine or fresh water) having an organ-system level of organisation. They are bilaterally symmetrical, triploblastic and coelomate animals. Structure: Body is covered by a calcareous shell and is unsegmented with a distinct head, muscular foot and visceral hump. A soft and spongy layer of skin forms a mantle over the visceral hump. The space between the hump and the mantle is called the mantle cavity in which feather like gills are present. They have respiratory and excretory functions. The anterior head region has sensory tentacles. The mouth contains a file-like rasping organ for feeding, called radula. They are usually dioecious and oviparous with indirect development. Examples: Pila (Apple snail), Pinctada (Pearl oyster), Sepia (Cuttlefish), Loligo (Squid), Octopus (Devil fish), Aplysia (Seahare), Dentalium (Tusk shell) and Chaetopleura (Chiton). Phylum – Echinodermata These animals have an endoskeleton of calcareous ossicles and, hence, the name Echinodermata (Spiny bodied). All are marine with organ-system level of organisation. The adult echinoderms are radially symmetrical but larvae are bilaterally symmetrical. They are triploblastic and coelomate animals. Digestive system is complete with mouth on the lower (ventral) side and anus on the upper (dorsal) side. The most distinctive feature of echinoderms is the presence of water vascular system which helps in locomotion, capture and transport of food and respiration. An excretory system is absent. Sexes are separate. Reproduction is sexual. Fertilisation is usually external. Development is indirect with free-swimming larva. Examples: Asterias (Star fish), Echinus (Sea urchin), Antedon (Sea lily), Cucumaria (Sea cucumber) and Ophiura (Brittle star). Phylum – Hemichordata Hemichordata was earlier considered as a sub-phylum under phylum Chordata. But now it is placed as a separate phylum under non-chordata. This phylum consists of a small group of worm-like marine animals with organ-system level of organisation. They are bilaterally symmetrical, triploblastic and coelomate animals. The body is cylindrical and is composed of an anterior proboscis, a collar and a long trunk. Circulatory system is of open type. Respiration takes place through gills. Excretory organ is proboscis gland. Sexes are separate. Fertilisation is external. Development is indirect. Examples: Balanoglossus and Saccoglossus. Phylum – Chordata Animals belonging to phylum Chordata are fundamentally characterised by the presence of a notochord, a dorsal hollow nerve cord and paired pharyngeal gill slits. These are bilaterally symmetrical, triploblastic, coelomate with organ-system level of organisation. They possess a post anal tail and a closed circulatory system. Phylum Chordata is divided into three subphyla: 1. Urochordata or Tunicata, 2. Cephalochordata and 3. Vertebrata. Subphyla Urochordata and Cephalochordata are often referred to as protochordates and are exclusively marine. In Urochordata, notochord is present only in larval tail, while in Cephalochordata, it extends from head to tail region and is persistent throughout their life. Examples: Urochordata – Ascidia, Salpa, Doliolum; Cephalochordata – Branchiostoma (Amphioxus or Lancelet). The members of subphylum Vertebrata possess notochord during the embryonic period. The notochord is replaced by a cartilaginous or bony vertebral column in the adult. Thus all vertebrates are chordates but all chordates are not vertebrates. Besides the basic chordate characters, vertebrates have a ventral muscular heart with two, three or four chambers, kidneys for excretion and osmoregulation and paired appendages which may be fins or limbs. Comparison of Chordates and Non-chordates Class – Cyclostomata All living members of the class Cyclostomata are ectoparasites on some fishes. They have an elongated body bearing 6-15 pairs of gill slits for respiration. Cyclostomes have a sucking and circular mouth without jaws. Their body is devoid of scales and paired fins. Cranium and vertebral column are cartilaginous. Circulation is of closed type. Cyclostomes are marine but migrate for spawning to fresh water. After spawning, within a few days, they die. Their larvae, after metamorphosis, return to the ocean. Examples: Petromyzon (Lamprey) and Myxine (Hagfish). Class – Chondrichthyes They are marine animals with streamlined body and have cartilaginous endoskeleton. Mouth is located ventrally. Notochord is persistent throughout life. Gill slits are separate and without operculum (gill cover). The skin is tough, containing minute placoid scales. Teeth are modified placoid scales which are backwardly directed. Their jaws are very powerful. These animals are predaceous. Due to the absence of air bladder, they have to swim constantly to avoid sinking. Heart is two-chambered (one auricle and one ventricle). Some of them have electric organs (e.g., Torpedo) and some possess poison sting (e.g., Trygon). They are cold-blooded (poikilothermous) animals, i.e., they lack the capacity to regulate their body temperature. Sexes are separate. In males pelvic fins bear claspers. They have internal fertilisation and many of them are viviparous. Examples: Scoliodon (Dog fish), Pristis (Saw fish), Carcharodon (Great white shark), Trygon (Sting ray). Class – Osteichthyes It includes both marine and fresh water fishes with bony endoskeleton. Their body is streamlined. Mouth is mostly terminal. They have four pairs of gills which are covered by an operculum on each side. Skin is covered with cycloid/ctenoid scales. Air bladder is present which regulates buoyancy. Heart is two-chambered (one auricle and one ventricle). They are cold-blooded animals. Sexes are separate. Fertilisation is usually external. They are mostly oviparous and development is direct. Examples: Marine – Exocoetus (Flying fish), Hippocampus (Sea horse); Freshwater – Labeo (Rohu), Catla (Katla), Clarias (Magur); Aquarium – Betta (Fighting fish), Pterophyllum (Angel fish). Class – Amphibia As the name indicates (Gr., Amphi : dual, bios, life), amphibians can live in aquatic as well as terrestrial habitats. Most of them have two pairs of limbs. Body is divisible into head and trunk. Tail may be present in some. The amphibian skin is moist (without scales). The eyes have eyelids. A tympanum represents the ear. Alimentary canal, urinary and reproductive tracts open into a common chamber called cloaca which opens to the exterior. Respiration is by gills, lungs and through skin. The heart is threechambered (two auricles and one ventricle). These are cold-blooded animals. Sexes are separate. Fertilisation is external. They are oviparous and development is direct or indirect. Examples: Bufo (Toad), Rana (Frog), Hyla (Tree frog), Salamandra (Salamander), Ichthyophis (Limbless amphibia). Class – Reptilia The class name refers to their creeping or crawling mode of locomotion (Latin, repere or reptum, to creep or crawl). They are mostly terrestrial animals and their body is covered by dry and cornified skin, epidermal scales or scutes. They do not have external ear openings. Tympanum represents ear. Limbs, when present, are two pairs. Heart is usually three-chambered, but four-chambered in crocodiles. Reptiles are poikilotherms (A poikilotherm is an animal whose internal temperature varies along with that of the ambient environmental temperature.). Snakes and lizards shed their scales as skin cast. Sexes are separate. Fertilisation is internal. They are oviparous and development is direct. Examples: Chelone (Turtle), Testudo (Tortoise), Chameleon (Tree lizard), Calotes (Garden lizard), Crocodilus (Crocodile), Alligator (Alligator). Hemidactylus (Wall lizard), Poisonous snakes – Naja (Cobra), Bangarus (Krait), Vipera (Viper). Class – Aves The characteristic features of Aves (birds) are the presence of feathers and most of them can fly except flightless birds (e.g., Ostrich). They possess beak. The forelimbs are modified into wings. The hind limbs generally have scales and are modified for walking, swimming or clasping the tree branches. Skin is dry without glands except the oil gland at the base of the tail. Endoskeleton is fully ossified (bony) and the long bones are hollow with air cavities (pneumatic). The digestive tract of birds has additional chambers, the crop and gizard. Endoskeleton is fully ossified (bony) and the long bones are hollow with air cavities (pneumatic). The digestive tract of birds has additional chambers, the crop and gizard. Heart is completely fourchambered. They are warm-blooded (homoiothermous) animals, i.e., they are able to maintain a constant body temperature. Respiration is by lungs. Air sacs connected to lungs supplement respiration. Sexes are separate. Fertilisation is internal. They are oviparous and development is direct. Examples: Corvus (Crow), Columba (Pigeon), Psittacula (Parrot), Struthio (Ostrich), Pavo (Peacock), Aptenodytes (Penguin), Neophron (Vulture). Class – Mammalia The most unique mammalian characteristic is the presence of milk producing glands (mammary glands) by which the young ones are nourished. They have two pairs of limbs, adapted for walking, running, climbing, burrowing, swimming or flying. The skin of mammals is unique in possessing hair. External ears or pinnae are present. Different types of teeth are present in the jaw. Heart is four-chambered. They are homoiothermous. Respiration is by lungs. Sexes are separate and fertilisation is internal. They are viviparous with few exceptions and development is direct. Examples: Oviparous-Ornithorhynchus (Platypus); Viviparous - Macropus (Kangaroo), Pteropus (Flying fox), Camelus (Camel), Macaca(Monkey), Rattus (Rat), Canis (Dog), Felis (Cat), Elephas (Elephant), Equus (Horse), Delphinus (Common dolphin), Balaenoptera (Blue whale), Panthera tigris (Tiger), Panthera leo (Lion). Distinct Characters of 4 classes of the Superclass Tetrapoda: 1. Amphibia: Capable of living on both land and water. Respiration through lungs as well as through skin and through gills. Cold blooded. 2. Reptilia: Creeping locomotion. Hard skin with scale like structure. Temperature of body varies as per the surrounding environment. 3. Aves: Bones are pneumatic or hollow to reduce weight. Additionally fore limbs are modified into wings to assist in flying. Warm blooded. 4. Mammal: Female feeds baby by producing milk through mammary glands. Hair present on body. External ears present. Warm Blooded

MORPHOLOGY OF FLOWERING PLANT

Morphology of Flowering Plants

THE ROOT

Tap Root System: In majority of the dicotyledonous plants, the direct elongation of the radicle leads to the formation of primary root which grows inside the soil. It bears lateral roots of several orders that are referred to as secondary, tertiary, etc. roots. The primary roots and its branches constitute the tap root system, as seen in the mustard plant.

Fibrous Root System: In monocotyledonous plants, the primary root is short lived and is replaced by a large number of roots. These roots originate from the base of the stem and constitute the fibrous root system, as seen in the wheat plant.

Adventitious Roots: In some plants, like grass, Monstera and the banyan tree, roots arise from parts of the plant other than the radicle and are called adventitious roots.

Functions of Roots: The main functions of the root system are absorption of water and minerals from the soil, providing a proper anchorage to the plant parts, storing reserve food material and synthesis of plant growth regulators.

Regions of the Root

The root is covered at the apex by a thimble-like structure called the root cap. It protects the tender apex of the root as it makes its way through the soil. A few millimetres above the root cap is the region of meristematic activity. The cells of this region are very small, thin-walled and with dense protoplasm. They divide repeatedly. The cells proximal to this region undergo rapid elongation and enlargement and are responsible for the growth of the root in length. This region is called the region of elongation. The cells of the elongation zone gradually differentiate and mature. Hence, this zone, proximal to region of elongation, is called the region of maturation. From this region some of the epidermal cells form very fine and delicate, thread-like structures called root hairs. These root hairs absorb water and minerals from the soil.

Modifications of Root

Roots in some plants change their shape and structure and become modified to perform functions other than absorption and conduction of water and minerals. They are modified for support storage of food and respiration.

Modification for Food Storage: Tap roots of carrot, turnips and adventitious roots of sweet potato, get swollen and store food.

Modification for Support: Banyan trees, after growing to a large size, develop hanging roots from branches. These are called prop roots. Similarly, the stems of maize and sugarcane have supporting roots coming out of the lower nodes of the stem. These are called stilt roots.

Modification for Respiration: In some plants such as Rhizophora growing in swampy areas, many roots come out of the ground and grow vertically upwards. Such roots, called pneumatophores, help to get oxygen for respiration.

THE STEM

The stem is the ascending part of the axis bearing branches, leaves, flowers and fruits. It develops from the plumule of the embryo of a germinating seed. The stem bears nodes and internodes. The region of the stem where leaves are born are called nodes while internodes are the portions between two nodes. The stem bears buds, which may be terminal or axillary. Stem is generally green when young and later often become woody and dark brown.

Functions of Stem: The main function of the stem is spreading out branches bearing leaves, flowers and fruits. It conducts water, minerals and photosynthates. Some stems perform the function of storage of food, support, protection and of vegetative propagation.

Modifications of Stem

Stems are modified to perform different functions. Underground stems of potato, ginger, turmeric, zaminkand, Colocasia are modified to store food in them. They also act as organs of perenation to tide over conditions unfavourable for growth. Stem tendrils which develop from axillary buds, are slender and spirally coiled and help plants to climb such as in gourds (cucumber, pumpkins, watermelon) and grapevines. Axillary buds of stems may also get modified into woody, straight and pointed thorns. Thorns are found in many plants such as Citrus, Bougainvillea. They protect plants from browsing animals. Some plants of arid regions modify their stems into flattened (Opuntia), or fleshy cylindrical (Euphorbia) structures. They contain chlorophyll and carry out photosynthesis. Underground stems of some plants such as grass and strawberry, etc., spread to new niches and when older parts die new plants are formed. In plants like mint and jasmine a slender lateral branch arises from the base of the main axis and after growing aerially for some time arch downwards to touch the ground. A lateral branch with short internodes and each node bearing a rosette of leaves and a tuft of roots is found in aquatic plants like Pistia and Eichhornia. In banana, pineapple and Chrysanthemum, the lateral branches originate from the basal and underground portion of the main stem, grow horizontally beneath the soil and then come out obliquely upward giving rise to leafy shoots.

THE LEAF

The leaf is a lateral, generally flattened structure borne on the stem. It develops at the node and bears a bud in its axil. The axillary bud later develops into a branch. Leaves originate from shoot apical meristems and are arranged in an acropetal order. They are the most important vegetative organs for photosynthesis.

A typical leaf consists of three main parts: leaf base, petiole and lamina. The leaf is attached to the stem by the leaf base and may bear two lateral small leaf like structures called stipules. In monocotyledons, the leaf base expands into a sheath covering the stem partially or wholly.

In some leguminous plants the leafbase may become swollen, which is called the pulvinus. The petiole help hold the blade to light. Long thin flexible petioles allow leaf blades to flutter in wind, thereby cooling the leaf and bringing fresh air to leaf surface.

The lamina or the leaf blade is the green expanded part of the leaf with veins and veinlets. There is, usually, a middle prominent vein, which is known as the midrib. Veins provide rigidity to the leaf blade and act as channels of transport for water, minerals and food materials. The shape, margin, apex, surface and extent of incision of lamina varies in different leaves.

Venation

The arrangement of veins and the veinlets in the lamina of leaf is termed as venation. When the veinlets form a network, the venation is termed as reticulate. When the veins run parallel to each other within a lamina, the venation is termed as parallel. Leaves of dicotyledonous plants generally possess reticulate venation, while parallel venation is the characteristic of most monocotyledons.

Types of Leaves

A leaf is said to be simple, when its lamina is entire or when incised, the incisions do not touch the midrib. When the incisions of the lamina reach up to the midrib breaking it into a number of leaflets, the leaf is called compound. A bud is present in the axil of petiole in both simple and compound leaves, but not in the axil of leaflets of the compound leaf.

The compound leaves may be of two types. In a pinnately compound leaf a number of leaflets are present on a common axis, the rachis, which represents the midrib of the leaf as in neem. In palmately compound leaves, the leaflets are attached at a common point, i.e., at the tip of petiole, as in silk cotton.

Phyllotaxy

Phyllotaxy is the pattern of arrangement of leaves on the stem or branch. This is usually of three types – alternate, opposite and whorled.

Alternate Arrangement of Leaves: In alternate type of phyllotaxy, a single leaf arises at each node in alternate manner, as in china rose, mustard and sun flower plants.

Opposite Arrangement of Leaves: In opposite type, a pair of leaves arise at each node and lie opposite to each other as in Calotropis and guava plants.

Whorled Arrangement of Leaves: If more than two leaves arise at a node and form a whorl, it is called whorled, as in Alstonia.

Modifications of Leaves

Leaves are often modified to perform functions other than photosynthesis.

Tendrils: They are converted into tendrils for climbing as in peas or into spines for defence as in cacti.

Bulb: The fleshy leaves of onion and garlic store food.

In some plants such as Australian acacia, the leaves are small and short-lived. The petioles in these plants expand, become green and synthesise food. Leaves of certain insectivorous plants such as pitcher plant, venus-fly trap are also modified leaves.

THE INFLORESCENCE

A flower is a modified shoot wherein the shoot apical meristem changes to floral meristem. Internodes do not elongate and the axis gets condensed. The apex produces different kinds of floral appendages laterally at successive nodes instead of leaves. When a shoot tip transforms into a flower, it is always solitary. The arrangement of flowers on the floral axis is termed as inflorescence. Depending on whether the apex gets converted into a flower or continues to grow, two major types of inflorescences are defined – racemose and cymose.

Racemose Inflorescence: In racemose type of inflorescences the main axis continues to grow, the flowers are borne laterally in an acropetal succession. Acropetal succession means that older flowers are at the base and younger flowers are near the apex.

Cymose Inflorescence: In cymose type of inflorescence the main axis terminates in a flower, hence is limited in growth. The flowers are borne in a basipetal order. In basipetal succession older flowers are at the apex and younger flowers are near the base.

THE FLOWER

The flower is the reproductive unit in the angiosperms. It is meant for sexual reproduction. A typical flower has four different kinds of whorls arranged successively on the swollen end of the stalk or pedicel, called thalamus or receptacle. These are calyx, corolla, androecium and gynoecium. Calyx and corolla are accessory organs, while androecium and gynoecium are reproductive organs. In some flowers like lily, the calyx and corolla are not distinct and are termed as perianth.

When a flower has both androecium and gynoecium, it is bisexual. A flower having either only stamens or only carpels is unisexual.

Symmetry of Flowers: In symmetry, the flower may be actinomorphic (radial symmetry) or zygomorphic (bilateral symmetry). When a flower can be divided into two equal radial halves in any radial plane passing through the centre, it is said to be actinomorphic, e.g., mustard, datura, chilli. When it can be divided into two similar halves only in one particular vertical plane, it is zygomorphic, e.g., pea, gulmohur, bean, Cassia. A flower is asymmetric (irregular) if it cannot be divided into two similar halves by any vertical plane passing through the centre, as in canna.

A flower may be trimerous, tetramerous or pentamerous when the floral appendages are in multiple of 3, 4 or 5, respectively. Flowers with bracts, reduced leaf found at the base of the pedicel, are called bracteate and those without bracts, ebracteate.

Hypogynous Flower: Based on the position of calyx, corolla and androecium in respect of the ovary on thalamus, the flowers are described as hypogynous perigynous and epigynous. In the hypogynous flower the gynoecium occupies the highest position while the other parts are situated below it. The ovary in such flowers is said to be superior, e.g., mustard, china rose and brinjal.

Perigynous Flower: If gynoecium is situated in the centre and other parts of the flower are located on the rim of the thalamus almost at the same level, it is called perigynous. The ovary here is said to be half inferior, e.g., plum, rose, peach.

Epigynous Flower: In epigynous flowers, the margin of thalamus grows upward enclosing the ovary completely and getting fused with it, the other parts of flower arise above the ovary. Hence, the ovary is said to be inferior as in flowers of guava and cucumber, and the ray florets of sunflower.

Parts of a Flower: Each flower normally has four floral whorls, viz., calyx, corolla, androecium and gynoecium.

Calyx: The calyx is the outermost whorl of the flower and the members are called sepals. Generally, sepals are green, leaf like and protect the flower in the bud stage. The calyx may be gamosepalous (sepals united) or polysepalous (sepals free).

Corolla: Corolla is composed of petals. Petals are usually brightly coloured to attract insects for pollination. Like calyx, corolla may be also free (gamopetalous) or united (polypetalous). The shape and colour of corolla vary greatly in plants. Corolla may be tubular, bell-shaped, funnel-shaped or wheel-shaped.

Aestivation: The mode of arrangement of sepals or petals in floral bud with respect to the other members of the same whorl is known as aestivation. The main types of aestivation are valvate, twisted, imbricate and vexillary.

Valvate: When sepals or petals in a whorl just touch one another at the margin, without overlapping, as in Calotropis, it is said to be valvate.

Twisted: If one margin of the appendage overlaps that of the next one and so on as in china rose, lady’s finger and cotton, it is called twisted.

Imbricate: If the margins of sepals or petals overlap one another but not in any particular direction as in Cassia and gulmohur, the aestivation is called imbricate.

Vexillary: In pea and bean flowers, there are five petals, the largest (standard) overlaps the two lateral petals (wings) which in turn overlap the two smallest anterior petals (keel); this type of aestivation is known as vexillary or papilionaceous.

Androecium

Androecium is composed of stamens. Each stamen which represents the male reproductive organ consists of a stalk or a filament and an anther. Each anther is usually bilobed and each lobe has two chambers, the pollen-sacs. The pollen grains are produced in pollen-sacs. A sterile stamen is called staminode. Stamens of flower may be united with other members such as petals or among themselves. When stamens are attached to the petals, they are epipetalous as in brinjal, or epiphyllous when attached to the perianth as in the flowers of lily. The stamens in a flower may either remain free (polyandrous) or may be united in varying degrees. The stamens may be united into one bunch or one bundle (monoadelphous) as in china rose, or two bundles (diadelphous) as in pea, or into more than two bundles (polyadelphous) as in citrus. There may be a variation in the length of filaments within a flower, as in Salvia and mustard.

Gynoecium

Gynoecium is the female reproductive part of the flower and is made up of one or more carpels. A carpel consists of three parts namely stigma, style and ovary. Ovary is the enlarged basal part, on which lies the elongated tube, the style. The style connects the ovary to the stigma. The stigma is usually at the tip of the style and is the receptive surface for pollen grains. Each ovary bears one or more ovules attached to a flattened, cushion-like placenta. When more than one carpel is present, they may be free (as in lotus and rose) and are called apocarpous. They are termed syncarpous when carpels are fused, as in mustard and tomato. After fertilisation, the ovules develop into seeds and the ovary matures into a fruit.

Placentation: The arrangement of ovules within the ovary is known as placentation. The placentation are of different types namely, marginal, axile, parietal, basal, central and free central.

Marginal: In marginal placentation the placenta forms a ridge along the ventral suture of the ovary and the ovules are borne on this ridge forming two rows, as in pea.

Axile: When the placenta is axial and the ovules are attached to it in a multilocular ovary, the placentaion is said to be axile, as in china rose, tomato and lemon.

Parietal: In parietal placentation, the ovules develop on the inner wall of the ovary or on peripheral part. Ovary is one-chambered but it becomes two-chambered due to the formation of the false septum, e.g., mustard and Argemone. When the ovules are borne on central axis and septa are absent, as in Dianthus and Primrose the placentation is called free central.

Basal: In basal placentation, the placenta develops at the base of ovary and a single ovule is attached to it, as in sunflower, marigold.

THE FRUIT

The fruit is a characteristic feature of the flowering plants. It is a mature or ripened ovary, developed after fertilisation. If a fruit is formed without fertilisation of the ovary, it is called a parthenocarpic fruit. Generally, the fruit consists of a wall or pericarp and seeds. The pericarp may be dry or fleshy. When pericarp is thick and fleshy, it is differentiated into the outer epicarp, the middle mesocarp and the inner endocarp.

In mango and coconut, the fruit is known as a drupe. They develop from monocarpellary superior ovaries and are one seeded. In mango the pericarp is well differentiated into an outer thin epicarp, a middle fleshy edible mesocarp and an inner stony hard endocarp. In coconut which is also a drupe, the mesocarp is fibrous.

THE SEED

The ovules after fertilisation, develop into seeds. A seed is made up of a seed coat and an embryo. The embryo is made up of a radicle, an embryonal axis and one (as in wheat, maize) or two cotyledons (as in gram and pea).

Structure of a Dicotyledonous Seed

The outermost covering of a seed is the seed coat. The seed coat has two layers, the outer testa and the inner tegmen. The hilum is a scar on the seed coat through which the developing seeds were attached to the fruit. Above the hilum is a small pore called the micropyle. Within the seed coat is the embryo, consisting of an embryonal axis and two cotyledons. The cotyledons are often fleshy and full of reserve food materials. At the two ends of the embryonal axis are present the radicle and the plumule. In some seeds such as castor the endosperm formed as a result of double fertilisation, is a food storing tissue. In plants such as bean, gram and pea, the endosperm is not present in mature seeds and such seeds are called nonendospermous.

Structure of Monocotyledonous Seed

Generally, monocotyledonous seeds are endospermic but some as in orchids are non-endospermic. In the seeds of cereals such as maize the seed coat is membranous and generally fused with the fruit wall. The endosperm is bulky and stores food. The outer covering of endosperm separates the embryo by a proteinous layer called aleurone layer. The embryo is small and situated in a groove at one end of the endosperm. It consists of one large and shield shaped cotyledon known as scutellum and a short axis with a plumule and a radicle. The plumule and radicle are enclosed in sheaths which are called coleoptile and coleorhiza respectively.

SEMI-TECHNICAL DESCRIPTION OF A TYPICAL FLOWERING PLANT

The description has to be brief, in a simple and scientific language and presented in a proper sequence. The plant is described beginning with its habit, vegetative characters – roots, stem and leaves and then floral characters inflorescence and flower parts.

Fusion is indicated by enclosing the figure within bracket and adhesion by a line drawn above the symbols of the floral parts. A floral diagram provides information about the number of parts of a flower, their arrangement and the relation they have with one another. The position of the mother axis with respect to the flower is represented by a dot on the top of the floral diagram. Calyx, corolla, androecium and gynoecium are drawn in successive whorls, calyx being the outermost and the gynoecium being in the centre.

DESCRIPTION OF SOME IMPORTANT FAMILIES

Fabaceae

This family was earlier called Papilonoideae, a subfamily of family Leguminosae. It is distributed all over the world.

Vegetative Characters: Trees, shrubs, herbs; root with root nodules

Stem: erect or climber

Leaves: alternate, pinnately compound or simple; leaf base, pulvinate; stipulate; venation reticulate.

Floral characters

Inflorescence: racemose

Flower: bisexual, zygomorphic

Calyx: sepals five, gamosepalous; imbricate aestivation

Corolla: petals five, polypetalous, papilionaceous, consisting of a posterior standard, two lateral wings, two anterior ones forming a keel (enclosing stamens and pistil), vexillary aestivation

Androecium: ten, diadelphous, anther dithecous

Gynoecium: ovary superior, mono carpellary, unilocular with many ovules, style single

Fruit: legume; seed: one to many, non-endospermic

Economic importance: Many plants belonging to the family are sources of pulses (gram, arhar, sem, moong, soyabean; edible oil (soyabean, groundnut); dye (indigofera); fibres (sunhemp); fodder (Sesbania, Trifolium), ornamentals (lupin, sweet pea); medicine (muliathi).

Solanaceae

It is a large family, commonly called as the ‘potato family’. It is widely distributed in tropics, subtropics and even temperate zones.

Vegetative Characters: Plants mostly, herbs, shrubs and small trees

Stem: herbaceous rarely woody, aerial; erect, cylindrical, branched, solid or hollow, hairy or glabrous, underground stem in potato (Solanum tuberosum)

Leaves: alternate, simple, rarely pinnately compound, exstipulate; venation reticulate

Floral Characters

Inflorescence : Solitary, axillary or cymose as in Solanum

Flower: bisexual, actinomorphic

Calyx: sepals five, united, persistent, valvate aestivation

Corolla: petals five, united; valvate aestivation

Androecium: stamens five, epipetalous

Gynoecium: bicarpellary, syncarpous; ovary superior, bilocular, placenta swollen with many ovules

Fruits: berry or capsule

Seeds: many, endospermous

Floral Formula

Economic Importance: Many plants belonging to this family are source of food (tomato, brinjal, potato), spice (chilli); medicine (belladonna, ashwagandha); fumigatory (tobacco); ornamentals (petunia).

Liliaceae

Commonly called the ‘Lily family’ is a characteristic representative of monocotyledonous plants. It is distributed world wide.

Vegetative characters: Perennial herbs with underground bulbs/corms/ rhizomes

Leaves: mostly basal, alternate, linear, exstipulate with parallel venation

Floral characters

Inflorescence: solitary / cymose; often umbellate clusters

Flower: bisexual; actinomorphic

Perianth: petal six (3+3), often united into tube; valvate aestivation

Androcium: stamen six, (3+3)

Gynoecium: tricarpellary, syncarpous, ovary superior, trilocular with many ovules; axile placentation

Fruit: capsule, rarely berry

Seed: endospermous

Economic Importance: Many plants belonging to this family are good ornamentals (tulip, Gloriosa), source of medicine (Aloe), vegetables (Asparagus), and colchicine (Colchicum autumnale).